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Meristems and Mitotic Cell Divisions
Meristems are the specific sites in the plant where mitotic divisions produce new cells. Most plants grow from the tips of branches and roots where apical meristems produce new cells. As a result, in all buds, at the ends of all branch tips and on root tips, apical meristems are found.
Apical meristems produce new cells that differentiate into three distinct embryonic tissues. These are the protoderm, ground meristem, and procambium. In the diagram, you can see protoderm will form the epidermal tissues. Ground meristem is the embryonic tissue that forms cells of the cortex and pith. Procambium forms the vascular tissues, xylem and phloem. Apical meristems contribute to initial plant growth called primary growth.
In stems and roots of woody plants, the procambium can persist to eventually form a continuous ring of meristematic tissue called the vascular cambium. This is the layer that produces new phloem and xylem tissues yearly, contributing to growth rings visible in woody plants. A second ring of meristematic tissue forms just below the epidermis in woody plants to produce bark. This is the cork cambium. The cork cambium and vascular cambium contribute to subsequent or secondary growth typically found only in woody plants
Now, roll over the apical meristem on the diagram..
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Apical Meristems in a Coleus Tip
In this 40x view of an apical bud of a coleus plant you can see dense masses of cells forming the apical meristems. Find the terminal bud with small leaf primordia on either side. Below the terminal bud are lateral or axillary buds at the base of larger leaf primordia. These buds also possess apical meristems. The dark lines in the leaves are procambium that will form the vascular tissues of the leaf midrib. The outer layer of cells on these buds is the protoderm. Now, roll over the right axillary bud to view an apical meristem of a root tip.
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Apical Meristem in a Root Tip
In root tips, apical meristems form cells needed for elongation and enlargement as the roots grow and penetrate through the soil. The meristem also forms a protective root cap. The cells to the upper left of this image are embryonic protoderm, ground meristem, and procambium.
Apical meristems are sites of intense mitotic cell divisions. The new cells produced fuel the growth and development of new tissues. At higher magnifications (400x to 1000x) cells of meristems can be seen in various stages of mitosis.
Now, let's take a look at cells of an apical meristem that are engaged in various stages of mitosis.
Forward to the mitosis module.
Back to the module outline.
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